Biography

Dr. R. Sanders Williams has served as a director of the Company since May 16, 2007. Dr. Williams serves currently as Senior Advisor for Science & Technology at Duke University and as President Emeritus of The J. David Gladstone Institutes, a non-profit biomedical research enterprise, holding both positions since January 1, 2018. Dr. Williams also currently is Professor of Medicine at the University of California San Francisco and Professor of Medicine at Duke University. Between February 1, 2021 and January 24, 2022 he fulfilled the role of Vice President (interim) for Research & Innovation at Duke University. Prior to these appointments he was president of The J. David Gladstone Institutes since November 2009, and he served as Chief Executive Officer of The J. David Gladstone Foundation, a non-profit organization supporting The J. David Gladstone Institutes, until December 31, 2018. Dr. Williams served Duke University between 2001 and 2010 as Dean of the School of Medicine, Senior Vice Chancellor, Senior Advisor for International Strategy, and founding Dean of the Duke-NUS Graduate Medical School Singapore. He has served previously as President of the Association of University Cardiologists, Chairman of the Research Committee of the American Heart Association, on the editorial boards of leading biomedical journals, on the Advisory Committee to the Director of the National Institutes of Health and on the board of external advisors of the National Heart, Lung and Blood Institute. Dr. Williams has served as a director of Tenaya Therapeutics, Inc. since October 2016 and has been a director of Amgen, Inc. since October 2014. Dr. Williams was a director of Bristol-Meyers Squibb Company from 2006 until 2013. Dr. Williams is a member of the National Academy of Medicine, and a Fellow of the American Association for the Advancement of Science.